Liguria
Description
The fossil collection, consisting of remains from various individuals discovered in caves in western Liguria, provides near complete evidence for the reconstruction of Ursus spelaeus, the cave bear, a species that was widespread throughout Europe and Asia during the Pleistocene.
The remains of this extinct bear are found with relative ease in the caves of western Liguria, from
the Savona area to the Imperia area. However, it is in the Toirano Caves that the highest concentration of remains is found, with truly spectacular accumulations. In particular, at the site known by the evocative name of "Bear Cemetery", two layers of accumulations have been identified. The first layer contains remains belonging to at least 139 individuals, including 57 cubs, 63 juveniles, and 19 adults, dating to a time span between 50.000 and 24.000 years ago. The second layer, however, has not yet been systematically investigated.
Analysis of the artefacts found in the so-called Sala dei Misteri (Hall of Mysteries), a section of the caves located further upstream, has revealed that the high concentration of bears is the result of a prolonged accumulation over time caused by physical phenomena. The remains, in fact, appear to have accumulated in this area after being transported from an external area directly connected to the underground environment.
Classification and size comparison
Kingdom: Animalia
Phylum: Chordata
Class: Mammalia
Order: Carnivora
Family: Ursidae
Genus: Ursus
Species: †Ursus spelaeus Rosenmüller, 1794
Modified from “Ursus spelaeus” evidence for the reconstruction of Hamelin de Guettelet, Wikimedia Commons, licenza CC BY-SA 3.0 (https://creativecommons.org/licenses/by-sa/3.0/).
Geologic Time Scale
- ERA: Cenozoic
- PERIOD: Quaternary
- EPOCH: Pleistocene
- ETÀ: Pleistocene Superiore (0.13-0.012 Ma)
Site of discovery
Various caves near Toirano (SV)
Display location
Toirano Caves (SV)
Catalogue number:
CdO1096
Gallery
A three-quarter view of the Ursus spelaeus on display at the Toirano Caves. Note the plantigrade posture, in which, as in primates, the entire palm of the hand and the sole of the foot rest on the ground during locomotion.
Courtesy of the Su gentile concessione del Ministero della Cultura, Soprintendenza Archeologia,
Belle Arti e Paesaggio for the provinces of Imperia and Savona.
Bibliography:
- Zunino, M., Starnini, E., Arobba, D., Avanzini, M., Citton, P., Firpo, M., … & Rellini, I. (2022). New insights into taphonomic analysis of the Upper Pleistocene Ursus spelaeus bone deposit from Bàsura cave (Toirano, NW Italy). Journal of Quaternary Science, 37(6), 1133-1147.
- Romano M., Citton P., Salvador I., Arobba D., Rellini I., Firpo M., Negrino F., Zunino M., Starnini E. & Avanzini M. (2019). A multidisciplinary approach to a unique palaeolithic human ichnological record from Italy (Bàsura Cave). eLife 2019, 8: e45204.
Credits and acknowledgments
We would like to extend our heartfelt thanks to Dr. Stefania Canavese of the Toirano Caves Service for sharing the multimedia material used on this page.
We would like to thank Dr. Stefano Costa and Dr. Marta Conventi of the Soprintendenza Archeologia Belle arti e Paesaggio for the provinces of Savona and Imperia for granting permission to use this material.
Reproduction, distribution, and any other use are prohibited without prior authorization from the aforementioned Superintendence.
We would also like to thank Professor Marta Zunino of the University of Genoa for her support and collaboration.